The interface pretends to be a thing. Leather has stitching, buttons have bevels, switches throw. You already know how it works because you have held one.
Definition
A skeuomorph is an ornamental feature carried over from an older material into a new one — the shutter click on a silent camera, the wood grain moulded into plastic. In software it means the screen imitates an object: light falls from above, surfaces have thickness, and pressing something moves it down and darkens it. Nothing here is decoration for its own sake. Every gradient is an argument that this rectangle is a physical control.
It reached its peak between 2007 and 2012, when a generation was meeting the touchscreen for the first time and needed to be told, in the most literal terms available, what could be poked.
The rules
Specimen
iOS 7 stripped it out in 2013 and the industry followed within eighteen months. The stated reason was maturity: users no longer needed a training-wheel metaphor. The unstated one was that stitched leather does not survive being resized to four hundred screen widths.
Anywhere the metaphor is load-bearing. Audio plug-ins still render knobs and VU meters because engineers reach for them the way they reach for hardware. Car dashboards, synthesisers, e-readers with page curl. The illusion earns its keep when muscle memory is the point.